1960 , William Llewellyn Brown , The Etruscan Lion , Yet the manelessness of the lioness is one of its most obvious features, and one which is never to my knowledge incorrectly shown by artists in the East where lions were familiar.
2002 , April , Philip Caputo , Maneless Lions , But are they really tougher? And if so, is there any connection between their manelessness and their ferocity?
2002 , July 9 , Anahad O'Connor , Shaggy, or Not So Shaggy: A New Look at Lions' Manes , All of these things led them to suspect that manelessness was typical in some parts of Kenya and might have been widespread in Africa many decades ago.