2007 , Victor Fet & ‎Alexi Popov , Biogeography and Ecology of Bulgaria , A total of 47 interstitial testate amoebas have been recorded in the Bulgarian Black Sea coast. Of these, 32 are psammobiotic, 11 are psammophilic, and 4 are psammoxenic.
2012 , H.U. Thiele , Carabid Beetles in Their Environments, Krogerus had already recognized that the influence of sand on psammophilic species is partly direct and partly indirect.
2016 , Rolf G. Beutel & ‎Richard A.B. Leschen , Coleoptera, Beetles. Morphology and Systematics, Most psammophilic species have expanded tibiae that function as shovels when these beetles are burrowing.